The extreme, workerless inquilines.

  • The extreme, workerless inquilines.

    Teleutomyrmex Kutter, 1950

    01) Teleutomyrmex schneideri Kutter, 1950
    (= Tetramorium schneideri (Kutter, 1950), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
    (not Tetramorium schneideri Emery, 1898)
    (= Tetramorium inquilinum Ward, Brady, Fisher & Schultz, 2015 ("2014"))
    02) Teleutomyrmex kutteri Tinaut, 1990
    (= Tetramorium kutteri (Tinaut, 1990), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
    (not Tetramorium semilaeve kutteri Santschi, 1927)
    03) Teleutomyrmex seiferti Kiran & Karaman, in Kiran, et al. 2017
    (= Tetramorium seiferti (Kiran & Karaman, in Kiran, et al. 2017), by analogy)
    04) Teleutomyrmex buschingeri Lapeva-Gjonova, in Kiran, et al. 2017
    (= Tetramorium buschingeri (Lapeva-Gjonova, in Kiran, et al. 2017), by analogy)

    Not yet described species of extreme, workerless inquiline, from the genus Teleutomyrmex Kutter, 1950

    05) The new, undescribed species from Teleutomyrmex Kutter, 1950 from Farab, Turkmenistan… See Dlussky, et al. 1990.

    Anergates Forel, 1874

    06) Anergates atratulus (Schenck, 1852), by Forel, 1874
    (= Myrmica atratula Schenck, 1852)
    [Also described as new by Schenck, 1853b]
    (= Tetramorium atratulum (Schenck, 1852), by Mayr, 1855)
    [= Tomognathus atratulus (Schenck, 1852), by Mayr, 1863 following Mayr, 1861, obsolete combination.]
    (= Tetramorium atratulum (Schenck, 1852), by Ward, Brady, Fisher & Schultz, 2015 ("2014"))
    07) Anergates friedlandi Creighton, 1934
    [= Tetramorium friedlandi (Creighton, 1934), by analogy]

    Tetramorium Mayr, 1855
    (Only a few species in a big genus.)

    08) Tetramorium microgyna Santschi, 1918
    09) Tetramorium parasiticum Bolton, 1980

    Pheidole Westwood, 1839
    (Only a few species in a big genus.)

    10) Pheidole neokohli Wilson, 1984
    (= Pheidole kohli (Wasmann, 1915), by Wilson, 1984)
    (= Anergatides kohli Wasmann, 1915)
    (not Pheidole kohli Mayr, 1901)
    11) Pheidole acutidens (Santschi, 1922), by Wilson, 1984
    (= Bruchomyrma acutidens Santschi, 1922)
    12) Pheidole argentina (Bruch, 1932), by Wilson, 1984
    (= Gallardomyrma argentina Bruch, 1932)
    13) Pheidole parasitica Wilson, 1984

    Excluded from the extreme, workerless inquilines
    Once this species was included in the extreme, workerless inquilines but now it is considered to be a workerless inquiline without extreme reductions, e.g. no pupoid males but normal ones. The decision to exclude it was made by Edward Osborne Wilson in 1984 in a study of the inquilines in the genus Pheidole Westwood, 1839.

    Pheidole Westwood, 1839
    (Only one species in a big genus.)

    14) Pheidole kusnezovi Wilson, 2003
    (= Pheidole symbiotica (Kusnezov, 1952), by Wilson, 1984)
    (= Eriopheidole symbiotica Kusnezov, 1952)
    (not Pheidole symbiotica Wasmann, 1909)

    Distribution

    01) Europe (Alps, Pyrenees and Northern Spain)
    02) Europe (Southern Iberia)
    03) Turkey (Anatolia)
    04) Europe (Southern Balkans)

    05) Turkmenistan

    06) Europe
    07) North America

    08) Southern Africa
    09) Southern Africa

    10) Central Africa
    11) South America
    12) South America
    13) India

    14) South America

    Host species

    01), 02), 03), 04), 05), 06), 07), 08) and 09) Certain species of the genus Tetramorium Mayr, 1855

    01) T. alpestre Steiner, Schlick-Steiner & Seifert, 2010 and T. impurum (Förster, 1850)
    and maybe T. caespitum (Linnaeus, 1758)?
    02) T. cf. caespitum (Linnaeus, 1758)
    03) T. cf. chefketi Forel, 1911
    04) T. cf. chefketi Forel, 1911

    05) A species from the genus Tetramorium Mayr, 1855…

    06) T. impurum (Förster, 1850), T. caespitum (Linnaeus, 1758), T. immigrans Santschi, 1927
    and T. moravicum Kratochvil, 1941, T. diomedeum Emery, 1908, T. chefketi Forel, 1911
    07) T. immigrans Santschi, 1927

    08) T. sericeiventre Emery, 1877 and T. sepositum Santschi, 1918
    09) T. avium Bolton, 1980

    10), 11), 12), 13) and 14) Certain species of the genus Pheidole Westwood, 1839

    10) P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912
    11) P. strobeli Emery, 1906
    12) P. nitidula Emery, 1888
    13) P. indica Mayr, 1879

    14) P. obscurior Forel, 1886

    A remark about synonymy

    Anergates friedlandi Creighton, 1934 is now a synonym from Anergates atratulus (Schenck, 1852), more precisely an introduced form in North Americe. So, the name is Anergates atratulus (Schenck, 1852) or, if you follow Ward et al. 2015 ("2014"), Tetramorium atratulum (Schenck, 1852)...

    Synonyms of host species

    - T. caespitum (Linnaeus, 1758) (= Formica caespitum Linnaeus, 1758)
    - T. impurum (Förster, 1850) (= Myrmica impura Förster, 1850)
    - P. megacephala (Fabricius, 1793) (= Formica megecephala Fabricius, 1793)
    - P. megacephala (Fabricius, 1793) subsp. melancholica Santschi, 1912 was originally described as P. punctulata Mayr, 1866 st. melancholica Santschi, 1912
    - The host species P. strobeli Emery, 1906 was, in 1922, described as a subspecies of an other species, nl. P. nitidula Emery, 1888 subsp. richteri Forel, 1909 (a much used junior synonym of the older P. perversa Forel, 1908 subsp. richteri Forel, 1909) and known as P. strobeli Emery, 1906 subsp. richteri Forel, 1909 since Santschi, 1916. The species P. strobeli Emery, 1906 was revived from synonymy (with P. nitidula Emery, 1888 since Wilson, 2003) and the subspecies was synonymized with it in Casadei-Ferreira et al., 2020.
    - P. obscurior Forel, 1886 was revived from synonymy with the 1 page older P. susannae Forel, 1886 and this in Casadei-Ferreira et al., 2020.

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